S W P S G A V C T T E T A S E S L K C S S S O H
Ludexv2026.11.01-65114139Dominant Species: MarineOfficial rulebook ↗

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Evolution and Domination

Dominant Species: Marine Evolution action (tile scoring and evolution cards) and the Domination action that claims special pawns.

Evolution

The Evolution row: card slots 1–5 above the terrain markers

Two steps: score one tile, then (perhaps) resolve an evolution card. Each of the five spaces shows a card-slot number (1–5) and a terrain marker terrain marker.

Evolution scoring

Choose exactly one tile of the terrain on your space’s marker and score it:

  1. Most species there: first value of the Tile Scoring table.
  2. Not a Vent: second most species gets the second value. An animal alone can’t take first and second.
  3. Land, Kelp, Reef or Seamount: third most gets the third value (it must be a third animal).
  4. Land with a fourth animal present: that animal gets 1 VP.

Ties on the number of species go to the animal higher on the food chain (Reptiles, then Cephalopods, then Fishes). An animal with no species on the tile scores nothing there.

Tile scoring table

Tile1st2nd3rd4th
Land Land8421
Kelp Forest Kelp Forest742—
Coral Reef Coral Reef632—
Seamount Seamount532—
Seagrass Meadow Seagrass Meadow42——
Sand Plain Sand Plain32——
Open Ocean Open Ocean21——
Hydrothermal Vent Hydrothermal Vent1———

Evolution example: a Kelp Forest with four animals’ cubes

Evolution card play

After scoring, if you have at least one thriving species on that tile, you must pick a face-up card from a slot numbered at or below your space’s number (space 5 = any card, space 1 = only slot 1).

Domination

The Domination row: the second space needs 3+ players, the third 4 players

Choose one element type you currently dominate (Dominance), then:

  1. Take control of its special pawn. If it is in front of a player or still on its setup space, put it in front of you, available. If it is on the action display, leave it there; you collect it when you next retrieve.
  2. Take its control marker control marker.
  3. Move its target marker target marker up the VP track to your domination value.

The middle space is open with 3 or more players, the right space only with 4.

PlayersDomination spaces
21
32
43
Rules

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